Position a baking rack in upper third of the oven; preheat oven to 400°F.
Line a 17.5 x 11.75-inch baking sheet with aluminum foil.
Place a 16.5 x 11.5-inch wire cooling rack inside the baking sheet.
Mist rack with cooking spray; set aside.
In a shallow pan, combine bread crumbs, 1/2 cup Parmesan cheese, 3 tablespoons parsley, paprika, and 1 teaspoon salt; mix well; set aside.
In a separate bowl, combine egg substitute and remaining 1 teaspoon salt, black pepper, marjoram, garlic powder, and oregano; mix well.
Dip both sides of mushrooms into egg mixture.
Dredge in crumb mixture until both sides are completely coated.
Transfer, stem side down, to the prepared wire rack.
Spray tops with cooking spray and bake for 15 to 20 minutes, or until mushrooms are tender.
Remove pan from oven.
Reset oven thermostat to 450°F.
Meanwhile, carefully turn mushrooms stem side up.
Cover each mushroom with 2 tablespoons of remaining Parmesan cheese, 1/4 of mozzarella cheese, and 1/4 cup tomato sauce; spray tops lightly.
Return to oven and bake until cheese melts and mushrooms are warmed through; about 10 minutes.
Remove from oven and use a wide spatula to transfer mushrooms to a serving platter.
Sprinkle with reserved 1 tablespoon parsley.
